Villy Christensen

Prospective Graduate Students / Postdocs

This faculty member is currently not looking for graduate students or Postdoctoral Fellows. Please do not contact the faculty member with any such requests.

Professor

Research Interests

Ecosystem modelling

Relevant Thesis-Based Degree Programs

Research Options

I am available and interested in collaborations (e.g. clusters, grants).
I am interested in and conduct interdisciplinary research.

Research Methodology

Ecopath with Ecosim ecosystem modelling framework

Great Supervisor Week Mentions

Each year graduate students are encouraged to give kudos to their supervisors through social media and our website as part of #GreatSupervisorWeek. Below are students who mentioned this supervisor since the initiative was started in 2017.

Villy has always given me the freedom and encouragement to work on a broad set of topics beyond my thesis work, and I am trully thankful for his helpful guidance.

Abdulrahman Ben Hasan (2018)

 

Graduate Student Supervision

Doctoral Student Supervision

Dissertations completed in 2010 or later are listed below. Please note that there is a 6-12 month delay to add the latest dissertations.

Population dynamics and interactions of sockeye and kokanee salmon (Oncorhynchus nerka) with piscivorous char (Salvelinus sp.) and rainbow trout (O. mykiss) in the large lakes and reservoirs of British Columbia (2022)

Freshwater fisheries management is focused on a combination of economic, social and conservation objectives. One of the trade-offs among these objectives involves a balance between harvest and consistent ecosystem functions. There can be conflicts between recreational fisheries and conservation, which are exacerbated by other habitat uses, such as dam operations. This dissertation examines the tools used by managers; how strategies employed by managers interact with natural systems and each other; what are some of the unexpected consequences; and what impact these tools have on recreational freshwater fisheries objectives. First, the use of nutrient addition as a tool to increase fish production is examined. Ecosystem modelling is then used to look at the impact of nutrient addition on target fish species in combination with different flow regimes in a highly managed reservoir. A decision analysis table is used to compare effects of different nutrient addition scenarios under variable flow conditions. Nutrient addition was found to be the most important driver of the system, with constant nutrient addition resulting in higher biomasses of the target fish species. Next, the outcomes of different stocking programs in British Columbia are summarized. The potential effect of salmon reintroduction programs on native fish species was predicted using ecosystem modelling. These models indicated that higher stocking levels had negative effects on the native kokanee salmon, but resulted in variable effects on the predators, depending on the species being stocked. Finally, ecosystem modelling is used to look at the possible effects of invasive non-native mussels in a productive lake. The model predicted that fish species that remained in the lake would be affected the most, while the invasive mussels had less of an effect on anadromous sockeye salmon and rainbow trout that reared in streams. This dissertation shows that maintaining consistent ecosystem functions to support recreational fisheries objectives can be challenge in itself.

On sustainability and financial return of fishery resources (2021)

Overexploitation and resource rent dissipation are some of the fundamental issues in fisheries management. The first undermines food security while the second implies a minimal economic return to the owning society. Sustainable fisheries are predominantly attained in conjunction with high management intensity, which keeps exploitation rates in check. Yet controlling exploitation becomes a daunting task under many complex fisheries contexts. Further, although profitability of fishing industry has improved after introducing quota-based systems, it is perceived that society is not receiving a fair share of the resource rent. In this dissertation, I focus on the Arabian Gulf region as a microcosm to examine various complex fisheries problems and underline, globally, the society鈥檚 compensation from the fishing industry. I begin by discussing situations where open access conditions are irreversible due to inherently poor management institutions or high dependency on fishing for livelihood. I show that well-designed size restriction鈥攁n easily implementable approach鈥攃an help avert overexploitation, rebuild depleted fish stocks and enhance yields without controlling exploitation rates. Next, I examine internationally shared fish stocks, whose sustainability requires managing exploitation rates at the international level rather than merely locally. I develop an age-structured model to evaluate bioeconomic trade-offs under alternative fishing scenarios. Harvesting a shared fish stock under cooperation or local but sustainable management provides much higher bioeconomic gains than competition. I then discuss the impacts of escalated market demand for dried swim bladder on fish, people and management in source countries. I highlight that while management interventions are required, the extremely high value of swim bladder would complicate regulatory efforts by stimulating black-market systems. Finally, I examine whether resource rent charges are imposed on catch share fisheries, and systematically compare that with forestry, oil, gas, and mining in 18 countries. I show that fishing is the only industry that consistently lacks resource rent charges, implying a forgoing stream of income in most countries. My dissertation contributes toward alleviating overfishing when exploitation rates are difficult to manage and underscores the need for national policies to consider the enhanced profitability of the fishing industry under catch share systems.

Addressing questions on the social and economic outcomes of an individual transferable quota fishery (2020)

In Canada, fisheries are expected to contribute to prosperous coastal communities and the maintenance of stable and viable fishing fleets, alongside other objectives that include conservation and complying with legal obligations to Indigenous Peoples. Individual transferable quotas (ITQs) have been promoted as a management approach to improve the conservation and economic outcomes of fisheries. The use of ITQs in British Columbia (BC) groundfish fisheries is widespread, following successive introductions of ITQs into the fisheries since 1990. There has been no comprehensive evaluation of the social and economic outcomes of ITQ management in the BC fisheries during this time, despite more than a decade of fishery participants and Indigenous and coastal community representatives raising concerns about the negative impacts of quota ownership and leasing. With a focus on the BC Pacific halibut fishery, I construct a database for licences and quota, including ownership and leasing. I examine changes in the ownership profile of the fishery over a 25-year period and consider the extent to which processors exercise control over the quota market through leasing. I construct a financial enterprise model based on accounting principles to assess the impact of quota ownership and leasing prices on the financial performance of owner-operator halibut vessels. I compare the results of this research against objectives for fisheries in Canada, determined through an extensive search of the literature, including current and historical policy and legal documents, conference proceedings, testimony to Senate and House of Commons committees, speeches and briefing material dating from the 1970s. Owner-operators have been increasingly marginalized in the halibut fishery. Owner-operators that have entered the fishery since 2001 catch 15% but own less than 1% of the halibut quota. Lease fees for halibut have regularly exceeded 80% of the landed price, reducing lessee fishing enterprises to minimal earnings that do not support reinvestment or renewal of the fleet. The BC halibut fishery is not meeting objectives for fisheries in Canada with respect to fleet viability and the equitable distribution of benefits. I provide an overview of measures that can be used for a just and fair transformation of fisheries to achieve socio-economic objectives.

Understanding, modeling and predicting trophic interactions between marine species (2015)

In this thesis, I explore and propose new methodologies and intend to provide new insights for understanding, modeling and predicting trophic interactions between marine species. I introduce the importance of trophic interactions in marine ecosystems and the common ecosystem modeling approaches applied to marine food webs. First, I present an attempt to synthetize information from all published Ecopath with Ecosim (EwE) models worldwide, gathered in the EcoBase digital repository. Through the compilation and standardization of a suite of metadata, I describe and discuss the usage of the EwE modeling approach and its evolution over time, since its very first application. I also present a meta-analysis of Ecopath models based on these metadata, where I select the models of potential interest, using a scoring method, and focus on one particular aspect of food web modeling, relating to the identification of keystone species. I propose a comprehensive and critical review of the ill-defined concept of keystone species and argue for a restored, exclusive and operational definition of the concept. The proposed definition is placed in a larger framework that considers different categories of ecologically important species. Then, a new functional index of keystoneness is derived from the EwE modeling approach, so as to identify potential keystone species in marine food webs. The proposed index addresses some of the biases observed in previously applied indices. Finally, I present an attempt to predict diet composition for predatory fish species. The intention is to build on existing large datasets, provided in the FishBase biodiversity information system, to identify predictors of fish feeding selectivity. The determination of clear or consistent pattern between biological and ecological species straits and diet composition is challenged by data restrictions, but some recommendations for future studies are provided. In conclusion, data availability may be a critical issue when considering some aspects of trophic interactions, especially for modeling and predictions at the species level. Data sharing within the scientific community, notably through the use of digital and open-access information repositories, is critical for the development of global meta-analyses in marine ecology.

Mortality of migrating Pacific salmon smolts in southern British Columbia, Canada (2010)

No abstract available.

Master's Student Supervision

Theses completed in 2010 or later are listed below. Please note that there is a 6-12 month delay to add the latest theses.

Impact of environmental variability on Jack Mackerel (Trachurus murphyi) spawning grounds in the open sea of the Southeast Pacific (2021)

Jack mackerel (Trachurus murphyi) is a migratory species broadly distributed in the Pacific Ocean. Its main spawning grounds are located in the high seas zone off the Chilean coast. Several ichthyoplankton surveys have been conducted in this region, showing shifts in egg distribution, which have become more noticeable in the last decade. It is widely known that environmental conditions play an important role in determining the optimal conditions for development and survival for early life stages. However, the environmental drivers underlying changes in the egg distribution have been little studied. Yet, this knowledge is critical for understanding the effects of environmental forcing on spawning grounds and trends in this fish鈥檚 population. The aim of this research was to elucidate how eggs distribution is mediated by the environment and determine the impacts of climate variability on spawning grounds. Egg presence data from 14 ichthyoplankton surveys (1999-2018), as well as remote sensing data, were used to model egg-environment relationships using the Maxent species distribution model. Specifically, I analyzed the optimum temperature parameter, tolerance range and maps of habitat suitability index. The results suggest that temperature is the main driver of egg distribution, and its interannual variability has an impact on eggs by reducing the occurrence probability due to abnormally warmer or cooler conditions. Further, the tolerance range defined a thermal strip that varied spatially in the N-S and E-W directions as a function of the sea surface temperature variability. The latitudinal shift of the egg distribution and the thermal strip were in sync. Thus, the northerly and southerly egg displacements may indicate the species' adaptive response to environmental fluctuations. Other factors, such as wind and temperature anomalies, also shaped suitable conditions for spawning. Changes in these parameters due to climate variability, resulted in unsuitable conditions and restricted egg distribution. Based on these findings, adaptive boundaries in the sampling design are recommended in future ichthyoplankton surveys to prevent sampling bias.

Fisheries in a changing environment: the impacts of the reduction in Shatt Al Arab flow on nearshore fish stocks in the Northern Persian Gulf (2017)

When fish catches decline, the standard recommended management solution is to reduce fishing mortality to allow stock recovery to more productive levels. This recommendation is based on the assumption that the most likely cause of the decline in the first place is fishing. Natural regime shifts and human-induced environmental changes are, however, often equally important factors in driving catch declines. In the Northern Persian Gulf, many commercial fish stocks are declining, raising questions about two main causes: overfishing and reduction in the flow of the major river, Shatt Al Arab. In Kuwait, the latter cause is strongly suspected of driving catch declines, especially with the implementation of high length limits and apparently good protection of juvenile nursery areas. Here I assess three case studies of Kuwait fish stocks and investigate the impact of reduced Shatt Al Arab flow on fish recruitment patterns. We found neutral and negative apparent capacity change in the green tiger shrimp stock and the orange-spotted grouper stock, respectively. These results suggest declining productivity in the nursery area of the orange-spotted grouper, but not in that of the tiger shrimp stock. In the case of the yellow-fin seabream assessment, the estimation of the relative recruitment was unreliable, hence the inability to examine the relationship between recruitment and the reduction in the flow rate of Shatt Al Arab. Our results demonstrate that reductions in Shatt Al Arab river flow are likely to impact fish recruitment patterns, causing changes in fish stock sizes. The findings presented here are expected to be a starting point for a more detailed investigation that tries to bring together data on what has been changing over time in the nearshore nursery environments, since most of the commercial fish stocks are inshore/estuarine dependent. Such investigation would be very critical for the fisheries management in deciding, for example, whether a reduction of fishing effort would be beneficial.

In search of "effective management": Case study of the British Columbia dungeness crab (Cancer magister) fisery and lessions from domestic and international experience (2013)

Canadian fisheries such as the British Columbia Dungeness crab (Cancer magister) fishery face a number of complex economic, political, and ecological challenges. The situation has been compounded by the increased complexity of management frameworks and declining resources of the government body entrusted with the management of Canada鈥檚 fishery resources. This thesis looks for best practices to create effective management within Canadian fisheries through investigation of three key components of fisheries management: Maintaining an exploited stock鈥檚 biological sustainability; developing and maintaining industry-government relationships; and providing equitable financing for management activities. Chapter one provides a comparison of management strategies within the British Columbia Dungeness crab and Western Australia rock lobster fisheries. This investigation shows that effective management to address the current challenges within the Dungeness crab fishery can be developed but will require time, resources, and building of a strong industry- government relationship. The second chapter outlines many of the challenges impacting Canadian fisheries, highlighting the recent impacts of the 鈥淟arocque Decision鈥. A recent resolution to the Larocque Decision (Bill C-38) is then explored utilizing an analysis of the joint project agreement for the snow crab fishery in Area 19 in eastern Canada. This chapter shows that collaborative management to address the fallout from the 鈥淟arocque Decision鈥 can be successfully developed under the current Canadian legislation. The chapter also demonstrates the limitations of current legislation in building such relationships and the fragility of collaborative frameworks built under the current regime. This thesis creates a list of the fundamental criteria needed to create effective management systems within fisheries, and concludes that collaborative management that creates a long-term true partnership between industry and government through a legislated process enhances the success of such management approaches.

Publications

  • (2021)
    Proceedings of the National Academy of Sciences, 118 (39)
  • (2021)
    Environmental Modelling & Software, 145, 105209
  • (2020)
    Frontiers in Marine Science, 7, 567877
  • (2020)
    Ecological Modelling, 428, 109087
  • (2020)
    Science, 368 (6496), 1243--1247
  • (2020)
    Reviews in Aquaculture, 12 (2), 1090--1100
  • (2020)
    Frontiers in Marine Science, 7, 554573
  • (2019)
    Ecological Modelling, 410, 108776
  • (2019)
    Lakes & Reservoirs: Research & Management, 8 (2), vii--9
  • (2019)
    Journal of Fish Biology,
  • (2019)
    Global Ecology and Conservation, , e00556
  • (2019)
    Global Ecology and Conservation, 17, e00556
  • (2018)
    Geoscientific Model Development, 11 (4), 1421--1442
  • (2018)
    Current Opinion in Environmental Sustainability, 29, 118--130
  • (2018)
    Journal of Environmental Radioactivity, 181, 118--127
  • (2018)
    Ocean and Coastal Management, 165, 33--38
  • (2018)
    Marine Pollution Bulletin, 129 (1), 1--7
  • Modelling marine trophic transfer of radiocarbon (14C) from a nuclear facility (2018)
    Environmental Modelling Software, 102, 138--154
  • (2017)
    Fish and Fisheries, 18, 40--53
  • (2017)
    Fish and Fisheries, 18 (1), 40--53
  • (2017)
    580, 17--36
  • Historical changes of the Mediterranean Sea ecosystem: modelling the role and impact of primary productivity and fisheries changes over time (2017)
    Nature Scientific Reports, 7, 44491
  • (2017)
    Fish and Fisheries, 19 (1), 1--29
  • (2017)
    Fish and Fisheries, 18 (4), 772--777
  • (2017)
    Coastal Management, 45 (1), 1--12
  • (2016)
    Ecological Modelling, 331, 173--184
  • (2016)
    319, 178--189
  • (2016)
    Ecological Modelling, 331, 142--150
  • Modelling the cumulative spatial鈥搕emporal effects of environmental drivers and fishing in a NW Mediterranean marine ecosystem (2016)
    Ecological Modelling, 331, 100--114
  • (2016)
    11 (5), e0155655
  • (2016)
    Ecological Modelling, 337, 272--280
  • (2015)
    312, 61--76
  • (2015)
    30 (11), 649--661
  • Evaluation of the effects on rockfish and kelp artisanal fisheries of the proposed Mejillones Peninsula marine protected area (northern Chile, SE Pacific coast) (2015)
    297, 141--153
  • Fishing Down through the Food Web (2015)
    40 (8), 370--372
  • (2015)
    Ecological Modelling, 302, 42--53
  • (2015)
    85 (1), 29--47
  • (2015)
    Reviews in Fish Biology and Fisheries, 25 (2), 413--424
  • (2015)
    533, 47--65
  • The global ocean is an ecosystem: simulating marine life and fisheries (2015)
    Global Ecology and Biogeography, 24, 507--517
  • (2015)
    313, 314--324
  • A framework to identify enabling and urgent actions for the 2020 Aichi Targets (2014)
    Basic and Applied Ecology, 15, 633--638
  • (2014)
    346 (6206), 241--244
  • (2014)
    143 (3)
  • (2014)
    Ecological Modelling, 294, 117--136
  • (2014)
    27, 19--31
  • Analysing changes in the southern Humboldt ecosystem for the period 1970鈥2004 by means of dynamic food web modelling (2014)
    Ecological Modelling, 274, 41--49
  • Animating the carbon cycle (2014)
    (17), 344--359
  • Assessing changes in the southern Humboldt in the 20th century using food web models (2014)
    278, 52--66
  • Fish biomass in the world ocean: a century of decline (2014)
    Marine Ecology Progress Series, 512, 155--166
  • (2014)
    9 (4), e95845
  • How sectors can contribute to sustainable use and conservation of biodiversity (2014)
  • (2014)
    Ecosystems, 17 (8), 1397--1412
  • Valuing seafood: The Peruvian fisheries sector (2014)
    Marine Policy, 44, 302--311
  • (2013)
    Ecological Modelling, 263, 139--151
  • EcoBase: a repository solution to gather and communicate information from EwE models (2013)
    (20)
  • (2013)
    38 (2), 76--81
  • Effects of hunting, fishing and climate change on the Hudson Bay marine ecosystem: I. Re-creating past changes 1970鈥2009 (2013)
    264, 130--142
  • Effects of hunting, fishing and climate change on the Hudson Bay marine ecosystem: II. Ecosystem model future projections (2013)
    264, 143--156
  • (2013)
    Deep-Sea Research Part II-Topical Studies in Oceanography, 75 (C), 1--15
  • Food-web structure of and fishing impacts on the Gulf of Cadiz ecosystem (South-western Spain) (2013)
    265, 26--44
  • (2013)
    BioScience, 63 (9), 735--744
  • (2013)
    Hydrobiologia, 717 (1), 85--108
  • Assessing the trophic position and ecological role of squids in marine ecosystems by means of food-web models (2012)
  • Ecopath Theory, Modeling, and Application to Coastal Ecosystems (2012)
    Treatise on Estuarine and Coastal Science, , 93--113
  • (2012)
  • (2012)
    Ecological Modelling, 228, 8--16
  • (2012)
    Marine Ecology Progress Series, 457, 261--264
  • (2012)
    102-1, 39--51
  • (2012)
    Fish and Fisheries, 13, 41--59
  • Global integrated assessment to support EU future environment policies (GLIMP) (2012)
  • Global integrated assessment to support EU future environment policies (GLIMP). Final report (2012)
  • (2012)
    7 (9), e43966
  • (2012)
    96 (2-3), 325--339
  • (2012)
    Global Ecology and Biogeography, 21 (4), 465--480
  • Trophic model of the outer continental shelf and upper slope demersal community of the southeastern Brazilian Bight (2012)
    76 (4), 763--779
  • (2011)
    Ecological Modelling, 222 (3), 857--864
  • Fishing down food webs (2011)
    Ecosystem Approaches to Fisheries: A Global Perspective, , 72--88
  • (2011)
    144, 2490--2498
  • Introduction: toward ecosystem-based management of fisheries (2011)
    Ecosystem Approaches to Fisheries: A Global Perspective, , 1--8
  • (2011)
    Marine Ecology Progress Series, 440, 255--266
  • (2011)
    Marine Ecology Progress Series, 442, 169--185
  • Predation on Fraser River sockeye salmon (2011)
  • (2011)
    Environmental Research, , 1--9
  • Progress in the use of ecosystem modeling for fisheries management (2011)
    Ecosystem Approaches to Fisheries: A Global Perspective, , 189--205
  • (2011)
    Ecological Modelling, 222 (9), 1569--1579
  • Scenario development for decision making (2011)
    Ecosystem Approaches to Fisheries: A Global Perspective, , 304--314
  • (2010)
    (15)
  • Behavior of sandeels feeding on herring larvae (2010)
    3, 164--168
  • Ecosystem structure and processes at Kaloko Honokhau, focusing on the role of herbivores, including the green sea turtle Chelonia mydas, in reef resilience (2010)
    Marine Ecology Progress Series, 420, 27--44
  • (2010)
    Global Change Biology, 16 (4), 1194--1212
  • (2010)
    221 (11), 1526--1544
  • MEY = MSY (2010)
    Fish and Fisheries, 11 (1), 105--110
  • Representation of multi-stanza life histories in Ecospace models for spatial organization of ecosystem trophic interaction patterns (2010)
    Bulletin of Marine Science, 86, 439--459
  • Using a fisheries ecosystem model with a water quality model to explore trophic and habitat impacts on a fisheries stock: A case study of the blue crab population in the Chesapeake Bay (2010)
    221, 997--1004
  • Using ecological indicators to assess the health of marine ecosystems: the North Atlantic (2010)
    CRC Handbook on Indicators of Ecosystem Health,
  • Constructing Historical Long-Term Primary Production in the Chesapeake Bay for Use in Ecosystem-Based Fisheries Management Models (2009)
  • (2009)
    Science, 323 (5912), 359--362
  • Database-driven models of the world's Large Marine Ecosystems (2009)
    Ecological Modelling, 220 (17), 1984--1996
  • Ecopath with Ecosim: Linking fisheries and ecology (2009)
    Handbook of Ecological Modelling and Informatics, , 55--70
  • (2009)
  • Life in Chesapeake Bay: an animated documentary (2009)
    53, 1--2
  • Mechanisms affecting recovery in an upwelling food web: The case of the southern Humboldt (2009)
    83 (1), 404--416
  • (2009)
    75 (7), 1773--1794
  • (2009)
    Ecological Modelling, 220 (20), 2802--2818
  • Policy options for the northern Benguela ecosystem using a multispecies, multifleet ecosystem model (2009)
  • (2009)
    Ecological Modelling, 220, 2583--2593
  • An Ecosim model for exploring ecosystem management options for the Gulf of Mexico: implications of including multistanza life history models for policy predictions (2008)
    Bulletin of Marine Science, 83 (1), 251--271
  • Description of the East Brazil Large Marine Ecosystem using a trophic model (2008)
    72 (3), 477--491
  • Fisheries in large marine ecosystems: descriptions and diagnoses (2008)
    The UNEP Large Marine Ecosystems Report: A Perspective on Changing Conditions in LMEs of the World's Regional Seas, , 23--40
  • Mercury in the Chesapeake (2008)
  • Models of the world's large marine ecosystems. GEF/LME global project Promoting Ecosystem-based Approaches to Fisheries Conservation and Large Marine \ldots (2008)
  • 脴kosystemmodel for Ringkobing Fjord. Skarvbestandens p\aavirkning af fiskebestandene (2008)
  • (2008)
    Canadian Journal of Fisheries and Aquatic Sciences, 65 (11), 2536--2551
  • Trophic flow kinetics in marine ecosystems: toward a theoretical approach to ecosystem functioning (2008)
    217, 33--47
  • Trophic modelling of the Peruvian upwelling ecosystem: Towards reconciliation of multiple datasets (2008)
    79 (2-4), 326--335
  • A synthesis of research activities at the Fisheries Centre on ecosystem-based fisheries modelling and assessment with emphasis on the Northern and Central Coast of BC (2007)
  • Adding realism to foraging arena predictions of trophic flow rates in Ecosim ecosystem models: shared foraging arenas and bout feeding (2007)
    209 (2-4), 342--350
  • Ecopath with Ecosim 6: the sequel (2007)
    43, 1--4 (September--October)
  • (2007)
    10 (3), 488--502
  • Riverine and early ocean migration and mortality patterns of juvenile steelhead trout (Oncorhynchus mykiss) from the Cheakamus River, British Columbia (2007)
    582, 55--65
  • Steller sea lions in the Gulf of Alaska: Modelling effects of environmental variation and fishing (2007)
    33, 14
  • Threats to the ocean: on the role of ecosystem approaches to fisheries (2007)
    46 (1), 67--86
  • A method for identifying keystone species in food web models (2006)
    195 (3-4), 153--171
  • Ecosystem modelling of dioxin distribution patterns in the marine environment (2006)
    On the multiple uses of forage fish: from ecosystems to markets, , 83--102
  • Ecosystem models show combined effects of fishing, predation, competition, and ocean productivity on Steller sea lions (\textitEumetopias jubatus) in Alaska (2006)
    Canadian Journal of Fisheries and Aquatic Sciences, 63, 2495--2517
  • Effects of fisheries on ecosystems: just another top predator? (2006)
    Top Predators in Marine Ecosystems, , 11--27
  • El futuro de las pesquerias (2006)
    Libro del Centenario de Chimbote,, , 339--345 and 402--403
  • Large whales and fisheries. Mapping top consumers in marine ecosystems. More food for thought (2006)
  • Turning the tide is easy enough (2006)
    Jan-F, 1--5
  • World fisheries: the next 50 years (2006)
    Science Magazine鈥檚 State of the Planet, 2006-2007, , 29--36
  • A global movement toward an ecosystem approach to management of marine resources (2005)
    Marine Ecology Progress Series, 300, 275--279
  • A stochastic framework for reconstructing historical marine mammal abundance from catch records and sparse abundance information: application to the Antarctic blue whale and North Atlantic and Arctic fin whale. (2005)
    C.M.2
  • Changes in the Gulf of Alaska ecosystems due to ocean climate change and fishing (2005)
  • Ecopath with Ecosim: a User's Guide, November 2005 Edition (2005)
  • Ecosystems and human wellbeing: Volume 1. Current state and trends (2005)
  • Food web models and data for studying fisheries and environmental impacts on Eastern Pacific ecosystems (2005)
  • Mapping global fisheries patterns and their consequences (2005)
    Place Matters 鈥 Geospatial Tools for Marine Science, Conservation and Management in the Pacific Northwest, , Part I--2
  • Possible ecosystem impacts of applying MSY policies from single-species assessment (2005)
    ICES Journal of Marine Science, 62 (3), 558--568
  • Quantitative ecosystem indicators for fisheries management (2005)
    ICES Journal of Marine Science, 62 (3), 307
  • Using ecosystem modeling for fisheries management: Where are we? (2005)
    , M:19
  • Automated parameter optimization for Ecopath ecosystem models (2004)
    172 (2-4), 141--149
  • Ecopath with Ecosim: a User's Guide, May 2004 Edition (2004)
  • (2004)
    Ecological Modelling, 172 (2-4), 109--139
  • Evaluating population trends of Steller sea lions in the Gulf of Alaska using ecosystem modelling (2004)
  • Modelling stock dynamics in the southern Benguela ecosystem for the period 1978-2002 (2004)
    African Journal of Marine Science, 26, 179--196
  • Opening the lost valley: implementing a 'Back to the Future' restoration policy for marine ecosystems and their fisheries (2004)
    Sustainable Management of North American Fisheries, , 165--193
  • Paris by day: Symposium on "Quantitative ecosystem indicators for fisheries management" (2004)
    12 (2), 4--6
  • (2004)
    172 (2-4), 103--107
  • Trade-offs in ecosystem-scale optimization of fisheries management policies (2004)
    Bulletin of Marine Science, 74 (3), 549--562
  • Trends in fish biomass off Northwest Africa, 1960-2000 (2004)
    Marine Fisheries, Ecosystems, and Societies in West Africa: Half a Century of Change, , 387--386
  • When South Africans swim together. Foreword to special issue "An ecosystem approach to fisheries management in the southern Benguela". (2004)
    African Journal of Marine Science, 26
  • Ecological geography as a framework for a transition toward responsible fishing (2003)
    Responsible Fisheries in the Marine Ecosystem, , 87--101
  • Fisheries impact on the South China Sea Large Marine Ecosystem: a preliminary analysis using spatially-explicit methodology (2003)
    Assessment, Management and Future Directions for Coastal Fisheries in Asian Countries, , 52--61
  • Hundred-year decline of North Atlantic predatory fishes (2003)
    Fish and Fisheries, 4 (1), 1--24
  • Mapping fisheries onto marine ecosystems for regional, oceanic and global integrations (2003)
    Trends in Exploitation, Protection, and Research, , 12:375--395
  • (2003)
    Science, 302 (5649), 1359--1361
  • Chapter 10: Ecosystem models (2002)
    Handbook of Fish and Fisheries, , 211--227
  • Towards sustainability in world fisheries (2002)
    418, 689--695
  • Coral reefs and other tropical fisheries (2001)
    Encyclopedia of Ocean Sciences, , 534--538
  • Ecosystems of the past: how can we know since we weren't there? (2001)
    Fisheries Centre Research Reports, 9 (4), 26--34
  • (2001)
  • Ecopath with Ecosim: a User's Guide, October 2000 Edition (2000)
  • Ecopath, Ecosim, and Ecospace as tools for evaluating ecosystem impact of fisheries (2000)
    ICES Journal of Marine Science, 57 (3), 697--706
  • Ecosystems of the North Atlantic: some definitions (2000)
    2, 1--4
  • Fishing down aquatic food webs (2000)
    88 (1), 46--51
  • (2000)
    51 (5), 447--450
  • Mapping fisheries onto marine ecosystems: a consensus approach for regional, oceanic and global integration (2000)
    , 16
  • Representing density dependent consequences of life history strategies in aquatic ecosystems: EcoSim II (2000)
    Ecosystems, 3 (1), 70--83
  • (1999)
    Ecosystems, 2 (6), 539--554
  • Ecosystem modelling and the challenge of ecosystem-based management (1999)
  • Proceedings of the EXPO '98 Conference on ocean food webs and economic productivity (1999)
  • Towards a stratified mass-balance model of trophic fluxes in the Gulf of Mexico Large marine ecosystem. (1999)
    The Gulf of Mexico Large Marine ecosystem, , 278--293
  • (1998)
    8 (1), S104--S109
  • Comparative modelling of trophic flows in four large upwelling ecosystems: global vs. local effects (1998)
    Global vs. Local Changes in Upwelling Ecosystems. Proc. of the first CEOS symposium, 5-9 September 1994, Monterey, CA,
  • Diet composition and trophic levels of marine mammals (1998)
    ICES Journal of Marine Science, 55 (3), 467--481
  • Fishery-induced changes in a marine ecosystem: insight from models of the Gulf of Thailand (1998)
    53, 128--142
  • Fishing down marine food webs (1998)
    279 (5352), 860--863
  • (1998)
    282 (5393), 1383--1383
  • How pervasive is "fishing down the food web" (1998)
    282 (5352), 1383a
  • The new Ecopath with Ecosim. (1998)
  • Competition between fisheries and marine mammals for prey and primary production in the Pacific Ocean (1997)
    Journal of Northwest Atlantic Fisheries Science, 22, 173--187
  • Flow modelling with ECOPATH: providing insights on the ecological state of agroecosystems (1997)
    Applications of Systems Approaches at the Farm and Regional Level, , 203--212
  • Placing fisheries resources in their ecosystem context (1997)
    10 (2), 9--14
  • Structuring dynamic models of exploited ecosystems from trophic mass-balance assessments (1997)
    Reviews in Fish Biology and Fisheries, 7 (2), 139--172
  • Ecological modeling for all (1996)
    19 (2), 25--26
  • (1996)
    6 (4), 417--442
  • Mass-balance models of northeastern Pacific ecosystems (1996)
  • Rehabilitating fished ecosystems: insights from the past (1996)
    19 (3), 13--14
  • (1996)
    6 (2), 243--247
  • A model of trophic interactions in the North Sea in 1981, the Year of the Stomach (1995)
    Dana, 11 (1), 1--28
  • A multispecies virtual population analysis incorporating information of size and age. (1995)
  • Coral reef systems and their exploitation: toward a global account. (1995)
  • (1995)
    Ecological Modelling, 72 (1-2), 129--144
  • Fish production, catches and the carrying capacity of the world oceans (1995)
    18 (3), 34--40
  • Primary production required to sustain global fisheries (1995)
    374 (6519), 255--257 [Erratum in Nature, 376: 279]
  • (1995)
    4 (3), 181--189.
  • Assessing trophic impacts of fish stock enhancement (1994)
  • Assessing trophic impacts of fish stock enhancement. Presented at the ICES Workshop to Evaluate the Potential for Stock Enhancement, 19-24 May, 1994, Charlottenlund, Denmark, 15 pp. (1994)
  • In our experience: a review of "Quantitative Fisheries Stock Assessment". (1994)
  • On the behavior of some proposed goal functions for ecosystem development (1994)
    75, 37--49
  • Reflections on the pelagic food web structure in the Black Sea (1994)
  • A trophic ecosystem model of Lake George, Uganda (1993)
    Trophic Models of Aquatic Ecosystems, , 124--129
  • An energy flow model of the mulberry dike-carp pond farming system of the Zhujiang Delta, Guangdong Province, China (1993)
    Trophic Models of Aquatic Ecosystems, , 48--55
  • Flow characteristics of aquatic ecosystems (1993)
    Trophic Models of Aquatic Ecosystems, , 338--352
  • Graphical representation of steady-state trophic ecosystem models (1993)
    Trophic models of aquatic ecosystems, , 20--28
  • On steady-state modeling of ecosystems (1993)
    Trophic Models of Aquatic Ecosystems, , 14--19
  • Steady-state modelling and network analysis. (1993)
  • Stratified models of large marine ecosystems: a general approach and an application to the South China Sea (1993)
    Large marine ecosystems: stress, mitigation and sustainability, , 148--174
  • A guide to the ECOPATH II program (version 2.1). ICLARM Software 6, 72 p. International Center for Living Aquatic Resources Management, Manila, Philippines. (1992)
  • A guide to the Ecopath II software system (version 2.1) (1992)
  • Ecological engineering and modelling: steady state modelling and network analysis. (1992)
  • ECOPATH II - An approach and a software for construction of ecosystem models and food web analysis. (1992)
  • (1992)
    Ecological Modelling, 61 (3-4), 169--185
  • Network analysis of trophic interactions in aquatic ecosystems. (1992)
  • On Gulland and Rosenberg (1992), a review. (1992)
  • Two new books on fish ecology from Chapman and Hall. Book review. (1992)
  • Una gu铆a al ECOPATH II sist茅ma de software (version 2.1). Traducido por Matthias Wolff. ZMT Contributions 1. Centro de Ecolog铆a Marina Tropical (ZMT), Bremen, Alemania, y Centro internac铆onal de Manejo de recursos vivos aquaticos (ICLARM), Manila, Filipin (1992)
  • A guide to the ECOPATH II software system (version 2.0) (1991)
  • Niche overlap - The predators add a dimension. (MS). (1991)
  • A draft guide to the ECOPATH II program (ver. 1.0). (1990)
  • Experimental estimation of food preference in cod 鈥 results of a pilot experiment (1990)
  • Global comparisons of aquatic ecosystems. (1990)
  • Larval growth and drift pattern and the separation of herring spawning groups in the North Sea (1990)
    37 (1), 135--148
  • Some features of fish food consumption estimates used by ecosystem modelers (1990)
  • The ECOPATH II model. (1990)
  • The ECOPATH II software or how we can gain from working together (1990)
    13 (2), 9--10
  • Estimation of larval growth rates from herring larvae surveys (1989)
  • HECAST a working group catch statistics spreadsheet package (1989)
  • Vertical migrations of herring, \emphClupea harengus, larvae in relation to light and prey distribution (1989)
    26 (2), 87--96
  • Plankton dynamics and larval herring growth, drift and survival in a frontal area (1988)
    Marine Ecology Progress Series, 44 (3), 205--219
  • Sampling of herring larvae close to a drifting buoy (1988)
  • Estimating average length composition from the International Herring Larvae Survey data (1986)
  • Herring larvae distribution, plankton production and hydrography in the Buchan area (Scotland) in September-October, 1985 (1986)
  • Studies of a larval herring (Clupea harengus L.) patch in the Buchan area. 2. Growth, mortality and drift of larvae (1986)
    6, 11--24
  • Estimation of herring larval production, (1985)
  • Improving the use of herring larval data in stock assessment. (1985)
  • Investigations on the relationship of herring larvae, plankton production and hydrography at Aberdeen Bank, Buchan Area, September 1984 (1985)
  • Summary of Danish enclosure experiments. (1985)
  • A combined fish larval, phytoplankton and oceanographic survey in the Skagerrak and the Kattegat in April 1983. (1983)
  • Predation by sand eel on herring larvae. (1983)
  • Sildens sideliniesystem. (1977)

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